Theory:

 
In this moral story, the narrator Prem, by narrating his own experience with a story he had told by an old man, shows us the importance of 'Unity'.
 
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Unity in diversity
  
The story reflects the importance of religious harmony, as, the author points out the fact of how everyone shares an equal role in the society irrespective of their colour, creed, race, religion, ranking, etc., and it would result in a huge disaster if this system is disturbed.
 
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Religious Harmony
 
Here, by inserting a story (story of Pambupatti) inside a story, the author tries to raise his voice against dictatorship, which is a lesson to the human beings themselves. It also reflects the advantages of unity and disadvantages of disunity. Though people belong to different categories, they all share the same position and deserve equal respect, importance, freedom, etc.
 
The narrator in this story, by portraying the character Makara, brings out how people suffer due to ineffective leadership and bad decisions. In this story, Makara being powerful, held the position of a leader and ruled the whole forest dominantly.
 
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Makara, the arrogant leader
 
It ruled so arrogantly in a way that it did not allow the fellow speakers to talk against its decision when expelling the other animals from the forest. It spotted out drawbacks in the other animals' behaviour and chased them one by one which includes tortoises, snakes, and lizards. These decisions taken by Makara had created trouble for itself and others as well, and so lost its power. Only when the animals had taken a group decision, the situation came back to normal at last. The story emphasizes the importance of being united.